(Not your issue....just FYI)
Propane Cylinders (not tanks) have a spring loaded shutoff valve in the OPD service valve.
If pigtail hose is not properly tightened it will NOT open the shutoff valve.
(Not your issue....just FYI)
The pigtail ACME Nut (Green) has the 'excess flow device'
IF OPD service valve is opened to quickly it will cause the ball bearing to move towards hose end and block flow thru hose.
Opening OPD service valve slowly allows pressure downstrean to equalize with upstream and ball bearing to move/remain centered allowing full flow thru hose
(Not your issue....just FYI)
The pigtail ACME Nut also has a 'thermal bushing' that will melt from high temp (fire). When bushing melts internal stub in ACME Nut moves away from OPD Service valve and it's spring loaded valve shuts stopping ANY flow out of cylinder
YOUR ISSUE
Because in service propane cylinder was NOT empty and you had NO leaks/excessive usage downstream....then problem is with regulator
Regulator either failed.......diaphragm cracked/leaked
OR
Regulator 'locked up' and would not allow propane to flow
If failed it will not properly function on either cylinder
If 'locked up' it will probably function just fine and be a 'fluke'
Replace regulator if you have another flow issue