In order to create air flow, the rooftop AC unit(s) must create pressure/vacuum. We could argue about that all day but lets not. No RV is air tight. There will always be air leaks of some type through vents in the roof/floor/slides, plumbing vents, the range hood vent, bath vents, dash AC vents (chassis AC), stateroom vents, partially open windows, etc. The vents WOULD NOT WORK if the RV was air tight. They would stall out. Think about it. Just the fact that moving air flows past an exterior vent will create some small pressure differential that will exchange air from inside to outside. It the leakage of air through these vents (leaks) that provides fresh air inside the RV.
If you still think an RV is air tight, drive one into a lake and see how long it floats!
To my knowledge, an RV rooftop AC unit differs from residential/commercial/industrial AC in that the latter is REQUIRED by the Uniform Mechanical Code to provide a certain % of fresh air to the recirc air. (usually 25% or less) Not so in vehicles. In vehicles is usually an option when/if provided.
Chum lee