I suspect the device at the bottom is the vapor recovery canister required on gasoline engines. This is your gas tank vent, And you should not be able to build vacuum on it.
Your control power (Vacuum) for the HEVAC system comes from a DIFFERENT line it may have a canister on it as well, but this is just a tank,, There will be two ports on it, one goes to the manifold and you should be able to suck, but not blow into it. The other port goes to the vacuum operated devices (HEVAC) and that one is open, no check valve. usually looks like a big ball, somewhere between softball and soccer ball in size, but can be shaped differently.
Home was where I park it. but alas the.
2005 Damon Intruder 377 Alas declared a total loss
after a semi "nicked" it. Still have the radios
Kenwood TS-2000, ICOM ID-5100, ID-51A+2, ID-880 REF030C most times